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An important issue to be considered when analysing time series is the assessment of the relative importance of different drivers. In the present case, only temperature and salinity were included in statistical models. It is however quite likely that other factors are also important, both for the production and distribution of planktonic species and where they may be distributed. Often different species may have overlapping or even complementary distributions and the distribution of one species may actually be a function of the presence of the other. This dependence could be described, for example, using a multivariate statistical approach [49], or even better with the use of species distribution models [50] taking into account potential interactions between the species and the environment. Furthermore, environmental drivers may also act synergistically on the production of planktonic species. In this case, a combination of approaches may be needed to detect synergistic effects. For example, for a given environmental variable, a multivariate statistical analysis could detect significant correlations between the abundance of two species, but a species distribution model could also detect the presence of a third species which is associated with that variable. In this case, the interaction between the two species could be tested by a negative binomial model in which the abundance of the two species is compared against the abundance of the third species. However, for the approach to be successful, more information on the species identification of each of the three species is needed.
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